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Deep in the Romanian forest, a team of scientists stumble upon the ruins of a 13th century abbey. On further inspection, they make a startling discovery: the abbey is built over the entrance to a giant underground cave system. These scientists are Dr. Nicolai and his associate Bacovia Radescu. Local biologists believe the cave could contain an undiscovered ecosystem, so they hire a group of American spelunkers to help them investigate its depths. Also tagging along is cameraman Alex Kim.
Jack and his brother Tyler are the thrill-seeking professional cave explorers hired by the scientists. The two run a team of the world famous divers. They arrive in Romania with the latest equipment, including a modified rebreather system allowing a diver to remain submerged for up to 24 hours. The diving team, which includes Charli, Briggs, sonar expert Strode, and Buchanan, begins its exploration. In the cave, they discover bizarre creatures, and also a group of explorers from three decades ago, who are no longer quite human.
The creatures within the movie are the result of a parasite from within the cave system. The parasite infects its human host and begins to transform it into a new humanoid built for surviving in cave systems. Several of the crew are infected with this parasite, including a woman who is able to escape in the end, supposedly with the intent to "infect the rest of the world".
